Dr. Holding did a Masters with Dr. Emily Taylor at Cal Poly, Ph.D. with Lisle Gibbs at Ohio State, and have postdoc'd with Chris Parkinson at Clemson, Darin Rokyta at Florida State, and Marjorie Matocq at UNR. I am currently a Research Investigator in the lab of David Ginsburg at University of Michigan. I have dedicated myself at each of these steps to the system that fascinates me most, pitvipers and their interactions with the environment. I focus most on the coevolutionary interactions between rattlesnakes and rodents, through the lens of venom toxicity and venom resistance. I combine genomics, proteomics, functional assays, and field ecology in my studies of these species' interactions.